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Rhinolophus ferrum-equinum (Neumann 1906). Rhinolophus eloquens (Loveridge ## # hus clivosus auger (= geoffroy-Tauger:) #: 1950B). R. sp. of Howard (TSUB)TequEIS R. CIRVOSus er = geoffroyi augér”) according to Bedford, (1932B). # clivosus Zambesiensis (Sudan record above).

Myotis (= Vespertilio) sp. (Neumann 1906). Myotis tricolor (seaf:#3: 9 rene caves material £io': SOUTH AFRICA above. Myotis macrodactylus (Arthur 1956A).

Miniopterus natalensis arenarius and Miniopterus schreibersi

(subsp. probably EEnrage) from Japan, M. EGEETEEi and M. E. schreibersi (Arthur #5.


Aside from the fact that larvae, nymphs, and females are taken

on bats, nothing is known concerning the biology of I. simplex. Males either do not take blood or feed very rapidly # secrete themselves thereafter; they should be searched for in retreats frequented by bats. Ixodes simplex is widely spread through the tropics and temperate climates o world and must be an uncommonly adaptable tick. Its hosts" ability to fly undoubtedly accounts in part for the great range of this species.




The haller's organ of both subspecies of I. simplex is like that of I. vespertilionis (Arthur 1956B).

*I am indebted to C. C. Sanborn, Curator of Mammals at Chicago Natural History Museum, and an outstanding authority on bats, for checking the bat host names in this section.


Females are readily separated from those of the only other known bat-infesting species of this genus, I. vespertilionis by the fact that I. simplex has normal-length legs, ugh the last pair is longer than usual (all pairs of legs of J. Vespertilionis are exceedingly long). Anal grooves are short and divergent. The scutum is slightly longer than broad, has gently curved posterolateral margins, and converging anterolateral margins; widely scattered, subequal punctations, shallow cervical grooves, no lateral grooves; its color is brownish, reddish, or yellowish. The basis capituli is triangular, without cornua or auriculae. Coxae are flat and without spurs. For a fuller description, see Arthur (1956A).

Males are unknown. Immature stages are described by Arthur (1956A).

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Oane - 567


English specimens.
(Natural History). Q loaned by Dr.


(Figures 232 to 235)


l TOrit Bat, unidentified Feb

This is the only specimen of 1. Vespertilionis known from the Sudan.


I. vespertilionis is widely distributed in the Old World and is known from scattered areas in Africa where search will probably reveal numerous new locality records.

The distribution of I. vespertilionis was first summarized by Nuttall and Warburton (1911) # Iater, more extensively, by Neumann (1916). The present distributional summary is based on the latter paper, with only subsequent reports added. More recently, Arthur (1956A) has brought these records up to date.


NORTH AFRICA: ALGERIA (Neumann 1916. Hirst 1916. Nuttall 1916. "Senevet 1937). MOROCCO (Arthur 1956A).

EAST AFRICA: SUDAN (Hoogstraal 1954B. Arthur 1956A).

UGANDA (Arthur 1956A). KENYA (HH collecting in crater of Mt. Menengai).

SOUTHERN AFRICA: UNION OF SOUTH AFRICA (Arthur 1956A. See NOTE five paragraphs below.

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GIBRALTAR (Neumann 1916). PORTUGAL (Hirst 1916). SPAIN Neumann 1916. Schulze 1927. Gil Collado 1936,1938,1948). FRANCE Neumann 1916. Hirst 1916. Jeannel 1926. Schulze l927. Senevet" 1937. Cooreman 1954A,B. Lamontellerie 1954. Arthur 1956A). GERMANY (Neumann 1916. Schulze 1923B,1944B. Schulze and Schlottke s. 1929). SWITZERLAND (Arthur 1956A). AUSTRIA (Neumann 1916. Nuttall, 1916). BRITISH ISLES (Neumann 1916. Nuttall 1916. Hirst 1916. * MacLeod 1939. Arthur 1948,1953A,1956A). BELGIUM (Bequaert 1913.” Schmitz and Bequaert 1914. Leruth 1939B. Cooreman 1951). LUXEMBURG (Leruth 1939B). NETHERLANDS (van Eyndhoven 1939,1953). TTALY (Neumann 1916. Tonelli-Rondelli 1930A). SARDINLA (Kohls, correspondence). GREECE (Schulze 1936. Pandazis 1947. Arthur 1956A). HUNGARY (Neumann 1916. Hirst 1916. Kotlan 1921A,B). CZECHOSLOWAKIA (Neumann 1910C. Rosicky 1953). BULGARIA (Schulze 1927). YUGOSLAVIA (Neumann 1916. Oswald 1939). ROMANIA (Leruth 1939A. Cooreman 1951). CRETE (Hirst 1916).

A NOTE: Schulze (1927) listed, a nymph from Rhinolophus hipposiderus (sic) (?ferrum-equinum) at "Zelebor" (?Europe).

Near East

TURKEY (Arthur 1956A). PALESTINE (Arthur 1956A). IRAN (Olenev 1927, 1931. Pomerantzev 1937,1950). RUSSIA (Olenev 1927, 1929, 1931. Pomerantzev 1937,1950. Karpov and Popov 1944).

Far East

JAPAN (From Sawada, Myiagi, Honshu, A. J. Nicholson legit: Kohls, correspondence).

NOTE: I. vespertilionis has been reported from Australia by Nuttall and #: and quoted by Neumann (1916), Ferguson (1925), and Leruth (1939B), but not subsequently verified. The host was listed as Ves o tricolor. The only vespertilionid bat known to have the specific name tricolor is # tricolor" of East and South Africa. The collecting locality for

*I am indebted to C. C. Sanborn, Curator of Mammals at Chicago Natural History Museum, and an outstanding authority on bats, for checking the bat host names in this section.

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