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Figure 328
SUDANESE FAUNAL AREAS BASED ON TICK
DISTRIBUTION
PLATE XVIC

[graphic]

Sudanese Faunal Areas Based On

Tick Distribution

A somewhat arbitrary, preliminary mapping of Faunistic Areas of tick distribution in the Sudan has been suggested (Figure 328, and pages 854 and 855). This scheme conforms as closely as permissable with that of Lewis (1953), which reflects the importance of critical and specialized factors of aquatic and semiaquatic breeding places of Tabanids as well as availability of hosts. Further study may reveal that additional divisions should be designated for ticks but the limits are not presently well understood. Lewis' Watershed, Marra, Nuba, Boma–Gemi, (western) Floodplain, Sudd, and Gezira Areas are not charted for ticks since data are not yet sufficient to indicate the specialized character of tick fauna in these regions. The more intensive tick collections from southeastern Sudan, however, allow the specification that Lewis' Torit-Loelli Area may be separated into two distinct areas, the Torit Area and the Kapoeta Area. Inasmuch as Lewis" western sector of the Floodplain Area is abandoned for the present purposes, the eastern sector is referred to here as the Eastern Floodplain Area.

WEST AFRICAN SUBREGION
UBANGLUELLE SAVANNAH DISTRICT
RAGA-LOKA AREA

The Raga-Loka Area, as here considered (Figure 328), is wider than that originally proposed by Chapin as the zone typical of West African fauna in the Sudan (Figure 1). It is a combination of Lewis' several southwestern Areas (Figure 327), the (Congo) Watershed, the Raga-Loka, the (western) Flood plain, and the Sudd. Consisting of various types of forests and savannahs with more numerous trees than found elsewhere in the Sudan, this Area largely lacks the broad grassy plains character of so much of the Sudan. Indicator tick species are those characteristic of the West African Subregion (page 852) though they represent only a small part of the total tick fauna endemic in West Africa. As already stated, indicator species typical of the West African Subregion are exceptional in the Sudan outside of the Raga-Loka Area.

Forests and wooded savannahs of this Area also shelter several species of the East and South African Subregion that are unknown elsewhere in the Sudan. "These are R. Sulcatus, R. Supertritus, R. tricuspis, and R. mihlensi. Restriction of these four ticks to this Area may indicate that "they are characteristic of more humid and shaded East African areas than those found elsewhere in the Sudan and therefore they have survived only here. Each is a species of little known biology.

EAST AFRICAN SUBREGION
EAST AFRICAN HIGHLAND DISTRICT
TORIT AREA

Data for this Area are more extensive than for any other in the Sudan. Commencing a few miles west of the Nile, which does not in itself appear to be an effective barrier, Torit Area is a vast, tree-dotted and clustered grassy plain that gradually becomes short grassy scrub east of Kapoeta. From its southern margin rise the various montane masses of limited extent that comprise the Imatong Area. To the north, the area becomes less densely wooded and numerous tick species vanish in the tall grass savannah. Few permanent streams break the plain but several extensive, poorly explored swamps fill depressions in the northern part of the Area.

Almost half of the tick species known from the Sudan have been collected in this Area. West African components are rare indeed owing primarily to the open forest or scattered tree aspect of the plains and absence of extensive forests fringing streams. As already noted, such moderately xerophilic species as A. lepidum and R. pravus occur in the eastern sector of this Area but are £ Iess common than in Kapoeta Area. Ticks requiring moderately cool temperatures and high humidity, such as R. appendiculatus, are absent here. Others that require moderate # such as A. variegatum, are common here but their numbers are not so great as when the same species occur in restricted situations in the Raga-Loka Area to the west.

IMATONG (MONTANE) AREA

The specialized tick fauna representative of both Subregions that is supported in these low, cool, partially forested mountains is listed on page 850.

The montane masses consist of the horseshoe shaped AcholiImatong Mountains, the highest peak of which is Mount Kinyeti (10,456 feet elevation). To the east, the Dongotona Mountains that rise to 8605 feet elevation have an isolated northwesterly extension in the Lafit Hills. The more eastern Didinga Hills with Mount Lotuke the highest point at 9169 feet elevation, are carried to the northwest by the Boya Hills. The base of these mountains in the plains is at an approximate altitude of 2000 feet. The vertebrate and tick fauna of none of these montane masses is well known.

SUDANESE SAWANNAH DISTRICT
KAPOETA AREA

Kapoeta Area, an extension of Karamoja District of Uganda, is characterized by three important species that are common here but become less numerous westwards toward Torit, their western boundary in southern Sudan. These are R. pravus, A. cohaerens, and A. lepidum. Only the last named species is known to extend northward into the Eastern Floodplain Area from where it fans out and becomes quite common in the Central Rainlands. A number of species present in adjacent Torit Area are not known from the Kapoeta Area. The more intensive work in the Torit Area may partially account for this disparity but the short grass scrub aspect of the eastern plains and the long and intense dry season are factors that may exclude many of these species. This is the area of vast migrations of tens of thousands of game animals from Upper Nile Province (Zaphiro 1949), a spectacular phenomenon that should be investigated for its significance in relation to tick distribution.

EASTERN FLOODPLAIN AREA

The picture of tick distribution in this Area is characterized by the presence of most of the widely ranging, common tropical species of the Sudan plains and by the apparent absence of numerous more specialized tropical species and northern species. More extensive exploration is required before the true composition of the tick fauna of this Area can be evaluated.

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